"Has improper value for" is a predefined property to track input errors for irregular value annotations that was likely caused by type or allowed value restrictions.
en:If a substrate association exists: part of the substrate which is affected in the interaction (e.g. 'leaves', if a fungus is growing on the leaves of an infected plant) +
If a substrate association exists: part of the substrate which is affected in the interaction (e.g. 'leaves', if a fungus is growing on the leaves of an infected plant) +